Estimating synonymous and nonsynonymous substitution rates under realistic evolutionary models. 引入突变和密码子偏见对这一方法进行了修正。
Comparing and Analysing on Models of Calculation and Statistical Testing of Nosynonymous Substitution Rate and Synonymous Substitution Rate during Gene Evolution 基因进化的同义与非同义替代计算及统计检验的比较分析